Journal Articles


Vimercati, G., Davies, S.J. & Measey, J. (in press) Rapid adaptive response to a mediterranean environment reduces phenotypic mismatch in a recent amphibian invader. Journal of Experimental Biology

Becker, FA, Tolley, KA, Measey, GJ & Altwegg, RA (2018) Extreme climate-induced life-history plasticity in an amphibian. American Naturalist 191(2): 250–258. pdf

Novoa, A., Shackleton, R., Canavan, S., Cybèle, C., Davies, S., Dehnen-Schmutz, K., Fried, J., Gaertner, M., Geerts, S., Griffiths, C., Kaplan, H., Kumschick, S., Le Maitre, D., Measey, J., Nunes, A.L., Richardson, D., Robinson, T.B., Touza, J. & Wilson, J.R.U (2018) A framework for engaging stakeholders on the management of alien species. Journal of Environmental Management 205: 286-297.


Nunes, A., Zengeya, T.A., Measey, J. & Weyl, O.L.F. (2017) Freshwater crayfish invasions in South Africa: past, present and potential future.  African Journal of Aquatic Science  42: 309-323. 5914.2017.1405788

De Villiers FA, Measey J. (2017Overland movement in African clawed frogs (Xenopus laevis): empirical dispersal data from within their native range. PeerJ 5:e4039

Vimercati, G., Davies, S.J., Hui, C. & Measey, J. (2017) Does restricted access limit management of invasive urban frogs? Biological Invasions19: 3659-3674.

Brun, C. M.A. Raquet, G.J. Measey & J.-M. Exbrayat (2017) Cyclic variation of the oviduct structure of Boulengerula taitana, an oviparous species of Gymnophiona: morphological changes, proliferation and apoptosis. African Journal of Herpetology66: 93-105. pdf

Courant, J., Vogt, S., de Villiers, A., Marques, R., Measey, J., Secondi, J., Rebelo, R., De Busschere, C., Ihlow, F., Backeljau, T., Rödder, D. & Herrel, A. (2017) Are invasive populations characterized by a broader diet than native populations? PeerJ5:e3250 DOI 10.7717/peerj.3250 pdf

Rödder D, Ihlow F, Courant J, Secondi J, Herrel A, Rebelo R, Measey GJ, Lillo F, de Villiers FA, De Busschere C, and Backeljau T (2017) Global realized niche divergence in the African-clawed frog Xenopus laevisEcology and Evolution 2017;00:115. pdf

Furman, B., Cauret, C., Colby, G., Measey, J. & Evans, B.J. (2017) Limited genomic consequences of hybridization between two African clawed frogs, Xenopus gilli and X. laevis (Anura: Pipidae). Scientific Reports7(1):1091 pdf

Vimercati, G., Davies, S.J., Hui, C. & Measey, J. (2017) Integrating age structured and landscape resistance models to disentangle invasion dynamics of a pond-breeding anuran. Ecological Modelling 356: 104–116 pdf

Vogt, S., de Villiers, F.A., Ihlow, F., Rödder, D. & Measey, J. (2017) Competition and feeding ecology in two sympatric Xenopus species (Anura: Pipidae). PeerJ5:e3130; DOI 10.7717/peerj.3130 pdf

Nunes, A., Zengeya, T.A., Hoffman, A.C., Measey, J. & Weyl, O.L.F. (2017) Distribution and establishment of the alien Australian redclaw crayfish, Cherax quadricarinatus, in South Africa and Swaziland. PeerJ5:e3135; DOI 10.7717/peerj.3135pdf

Kumschick, S., Measey, G.J., Vimercati, G., de Villiers, F.A., Mokhatla, M., Davies, S.J., Thorp, C.J., Rebelo, A., Blackburn, T. & Kraus, F. (2017) How repeatable is the Environmental Impact Classification of Alien Taxa (EICAT)? Comparing independent global impact assessments of amphibians. Ecology and Evolution 2017;00:110.

da Silva, J.M., Feldheim, K.A., Measey, G.J., Doucett-Riise, S., Daniels, R.J., Chauke, L.F. & Tolley, K.A. (2017) Genetic diversity and differentiation of the Western Leopard Toad (Sclerophrys pantherina) based on mitochondrial and microsatellite markers. African Journal of Herpetology 66: 25-38. pdf

Edwards, S, Tolley, KA & Measey, GJ (2017) Habitat characteristics influence the breeding of Rose's dwarf mountain toadlet Capensibufo rosei (Anura: Bufonidae) Herpetological Journal27: 287-298. pdf

Channing, A., Measey, G.J., De Villiers, A.L., Turner, A.A. & Tolley, K.A. (2017) Taxonomy of the Capensibufo rosei group (Anura: Bufonidae) from South Africa. Zootaxa 4232(2): 282–292 pdf

Kumschick, S., Vimercati, G., de Villiers, F.A., Mokhatla, M., Davies, S.J., Thorp, C.J., Rebelo, A. & Measey, G.J. (2017) Impact assessment with different scoring tools: How well do alien amphibian assessments match?  Neobiota 33:53-66pdf

Measey, J., Davies, S., Vimercati, G., Rebelo, A., Schmidt, W. & Turner, A.A. (2017) Invasive amphibians in southern Africa: a review of invasion pathways. Bothalia-African Biodiversity & Conservation  47(2), a2117. doi:10.4102/abc.v47i2.2117 pdf

Nunes, A., Hoffman, A.C., Zengeya, T.A., Measey, J. and Weyl, O.L.F. (2017) Red swamp crayfish, Procambarus clarkii, found in South Africa 22 years after attempted eradication. Aquatic Conservation: Marine and Freshwater Ecosystems  27: 1334–1340. pdf

Measey, G.J., Stevenson, B., Scott, T, Altwegg, R and Borchers, D. (2017) Counting chirps: acoustic monitoring of cryptic frogs. Journal of Applied Ecology 54: 894–902 pdf

Dollion, A., G.J. Measey, R. Cornette, L. Carne, K.A. Tolley, J. da Silva, R. Boistel, A-C. Fabre, and A. Herrel (2017) Does diet drive the evolution of head shape and bite force in chameleons of the genus Bradypodion? Functional Ecology 31: 671-684 pdf

Measey, J. (2017). Where do African clawed frogs come from? An analysis of trade in live Xenopus laevis imported into the USA. Salamandra53: 398-404. pdf


Measey, J. (2016) Overland movement in African clawed frogs (Xenopus laevis): a systematic systematic review. PeerJ  4:e2474

Measey, G.J., Vimercati, G., de Villiers, F.A., Mokhatla, M., Davies, S.J., Thorp, C.J., Rebelo, A. & Kumschick, S. (2016) A global assessment of alien amphibian impacts in a formal framework. Diversity and Distributions  22: 970-981. pdf

Herrel, A., C. Moureaux, M. Laurin, G. Daghfous, K. Crandell, K.A. Tolley, G.J. Measey, B. Vanhooydonck & R. Boistel (2016) Frog origins: inferences based on ancestral reconstructions of locomotor performance and anatomy. Fossil Imprint. 71: 239-248.

Matthews, T., Measey, G.J. & Roberts, D. (2016) Implications of a summer breeding frog from Langebaanweg (South Africa): regional climate evolution at 5.1 Mya. South African Journal of Science J Sci. 112(9/10): 20160070

Edwards, S., Herrel, A., Vanhooydonck, B., Measey, G.J. & Tolley, K.A. (2016) Diving in head first: Investigations into trade-offs between phenotypic traits and sand-diving predator escape strategy in desert lizards (Meroles, Lacertidae, Squamata). Biological Journal of the Linnean Society 119 (4): 919–931

Broadley, D.G. & Measey, G.J. (2016) A new species of Zygaspis (Reptilia: Squamata: Amphisbaenidae) from north-eastern Mozambique. African Journal of Herpetology 65(2): 115-122. pdf

Ihlow F, Courant J, Secondi J, Herrel A, Rebelo R, Measey GJ, Lillo F, de Villiers FA, Vogt S, De Busschere C, Backeljau T, and Rödder D. 2016. Impacts of climate change on the global invasion potential of the African clawed frog Xenopus laevisPLoS ONE 11(6): e0154869. doi:10.1371/journal.pone.0154869 pdf

Miranda, N.A.F., Measey, G.J., Peer, N., Raw, J.L., Perissinotto, R. & Appleton, C.C. (2016) Shell crushing resistance of alien and native thiarid gastropods to predatory crabs in South Africa. Aquatic Invasions 11: 303-311. pdf

Van Sittert, L. & Measey, G.J. (2016) Historical perspectives on global exports and research of African clawed frogs (Xenopus laevis). Transactions of the Royal Society of South Africa  71: 157-166.pdf

da Silva, J.M., Carne, L., Measey, G.J., Herrel, A. & Tolley, K.A. (2016) The relationship between diet, cranial morphology, bite performance and habitat in an adaptive radiation of dwarf chameleon (Bradypodion)? Biological Journal of the Linnean Society  119: 52-67. pdf

Janion-Scheepers, C., Measey, J., Braschler, B., Chown, S.L., Coetzee, L., Colville, J.F., Dames, J., Davies, A.B., Davies, S., Davis, A., Dippenaar-Schoeman, A.S., Duffy, G.A., Fourie, D., Griffiths, C., Haddad, C.R., Hamer, M., Herbert, D., Hugo-Coetzee, E.A., Jacobs, A., Jansen van Rensburg, C., Lamani, S., Lotz, L.N., Louw, S.vdM., Lyle, R., Malan, A.P., Marais, M., Neethling, J.-A., Nxele, T., Plisko, D., Prendini, L., Rink, A.N., Swart, A., Theron, P., Truter, M., Ueckermann, E., Uys, V.M., Villet, M.H., Willows-Munrow, S. & Wilson, J.R. (2016) Soil biota in a megadiverse country: current knowledge and future research directions in South Africa. Pedobiologia 59, 129-174. pdf

De Busschere, C., Courant, J., Herrel, A., Rebelo, R., Rödder, D., Measey, G.J. & Backeljau, T. (2016) Unequal contribution of native South African phylogeographic lineages to the invasion of the African clawed frog, Xenopus laevis, in Europe PeerJ 4:e1659 

de Villiers, F.A., de Kock, M. & de Measey, G.J. (2016) Controlling the African clawed frog Xenopus laevis to conserve the Cape platanna Xenopus gilli in South Africa. Conservation Evidence 13, 17. pdf

Kornijów, R., Measey, G.J. and Moss, B. (2016) The structure of the littoral: effects of waterlily density and perch predation on sediment and plant‐associated macroinvertebrate communities. Freshwater Biology 61,  32-50. pdf


Nxele, T.C., Lamani, S., Measey, G.J., Armstrong, A.J., Plisko, J.D., Willows-Munro, S., Janion-Scheepers, C. & Wilson, J.R.U. (2015). Studying earthworms (Annelida: Oligochaeta) in South Africa. African Invertebrates 56, 779-806. pdf

Raquet, M. Measey, G.J & Exbrayat, J.-M. (2015) Annual variation of ovarian structures of Boulengerula taitana (Loveridge 1935), a Kenyan caecilian. African Journal of Herpetology  64, 116-134. pdf

Junker, K., Mariaux, J., Measey, G.J. & Mutafchiev, Y. (2015) Meteterakis  saotomensis n. sp. (Nematoda: Heterakidae) from Schistometopum thomense (Bocage) (Gymnophiona: Dermophiidae) on São Tomé Island. Systematic Parasitology 92, 131–139 pdf 

Measey, G.J., Vimercati, G., de Villiers, F.A., Mokhatla, M.M., Davies, S.J., Edwards, S. & Altwegg, R. (2015). Frog eat frog: exploring variables influencing anurophagy. PeerJ  3, e1204.

Vanhooydonck B., Measey G.J., Edwards S., Makhubo B., Tolley K.A., & Herrel A. (2015). The effects of substrate on locomotor performance in lacertid lizards. Biological Journal of the Linnean Society 115, 869-881. pdf

Higham, T. E, Measey, G.J., Bim-Jeffery, A.V., Herrel, A. & Tolley K.A. (2015) Functional divergence between morphs of a dwarf chameleon: differential locomotor kinematics in relation to habitat structure. Biological Journal of the Linnean Society 116, 27-40. pdf

Cressey, E.R., Measey, G.J., & Tolley, K.A. (2015). Fading out of view: the enigmatic decline of Rose’s mountain toad Capensibufo rosei. Oryx, 49, 521–528. pdf

Mokhatla, M. M., Rödder, D. & Measey, G.J. (2015) Assessing the effects of climate change on distributions of Cape Floristic Region amphibians. South African Journal of Science;111, 2014-0389. pdf

Rhodin, A.G., Kaiser, H., van Dijk, P.P., Wüster, W., O’Shea, M., Archer, M., Auliya, M., Boitani, L., Bour, R., Clausnitzer, V., Contreras-MacBeath, T., Crother, B.I., Daza, J.M., Driscoll, C.A., Flores-Villela, O., Frazier, J., Fritz, U., Gardner, A.L., Gascon, C., Georges, A., Glaw, F., Grazziotin, F.G., Groves, C.P., Haszprunar, G., Havaš, P., Hero, J.-M., Hoffmann, M., Hoogmoed, M.S., Horne, B.D., Iverson, J.B., Jäch, M., Jenkins, C.L., Jenkins, R.K.B., Kiester, A.R., Keogh, J.S., Thomas E. Lacher, J., Lovich, J.E., Luiselli, L., Mahler, D.L., Mallon, D., Mast, R., McDiarmid, R.W., Measey, J., Mittermeier, R.A., Molur, S., Mosbrugger, V., Murphy, R.W., Naish, D., Niekisch, M., Ota, H., Parham, J.F., Parr, M.J., Pilcher, N.J., Pine, R.H., Rylands, A.B., Sanderson, J.G., Savage, J.M., Schleip, W., Scrocchi, G.J., Shaffer, H.B., Smith, E.N., Sprackland, R., Stuart, S.N., Vetter, H., Vitt, L.J., Waller, T., Webb, G., Wilson, E.O., Zaher, H. & Thomson, S. (2015) Comment on Spracklandus Hoser, 2009 (Reptilia, Serpentes, ELAPIDAE): request for confirmation of availability of the generic name and for the nomenclatural validation of the journal in which it was published (Case 3601; BZN 70: 234–237; 71: 30–38; 133-135,181-182, 252-253). Bulletin of Zoological Nomenclature, 72, 65-78. pdf

Conlon, J. M., Mechkarska, M., Coquet, L., Leprince, J., Jouenne, T., Vaudry, H., & Measey, G. J. 2015. Evidence from peptidomic analysis of skin secretions that allopatric populations of Xenopus gilli (Anura: Pipidae) constitute distinct lineages. Peptides. 63, 118-125. pdf

Stevenson, B.C., Borchers, D.L., Altwegg, R., Swift, R.J., Gillespie, D.M., Measey, G.J. (2015), A general framework for animal density estimation from acoustic detections across a fixed microphone array. Methods in Ecology and Evolution, 6, 38–48. pdf


Stoelting, R.J., Measey, G.J. & Drewes, R.C. (2014) Population genetics of the São Tomé caecilian (Gymnophiona: Dermophiidae: Schistometopum thomense) reveals strong geographic structuring. PLoS one, e104628. pdf

da Silva, J.M., Herrel, A., Measey, G.J., Vanhooydonck, B., & Tolley, K.A. (2014). Linking microhabitat structure, morphology and locomotor performance traits in a recent radiation of dwarf chameleons. Functional Ecology. 28, 702-713. pdf

Vanhooydonck, B., James, R.S., Tallis, J., Aerts, P., Tadic, Z., Tolley, K.A., Measey, G.J. & Herrel, A. (2014). Is the whole more than the sum of its parts? Evolutionary trade-offs between burst and sustained locomotion in lacertid lizards. Proceedingsof the Royal Society B: Biological Sciences281(1777), 20132677. pdf

da Silva, J.M., Herrel, A., Measey, G.J., & Tolley, K.A. (2014). Sexual Dimorphism in Bite Performance Drives Morphological Variation in Chameleons. PloSone9(1), e86846. pdf


Edwards, S., Tolley, K.A., Vanhooydonck, B., Measey, G.J. & Herrel, A. 2013. Is dietary niche breadth linked to morphology and performance in Sandveld lizards Nucras (Sauria: Lacertidae)? Biological Journal of the Linnean Society 110, 674-688. pdf

Segall, M., Tolley, K.A., Vanhooydonck, B., Measey, G.J. & Herrel, A. 2013. Impact of temperature on performance of South African Dwarf Chameleon, Bradypodion pumilum and B. occidentale? Journal of Experimental Biology 216, 3828-3836. pdf

Herrel, A., Tolley, K.A., Measey, G.J., da Silva, J.M., Potgieter, D.F., Boller, E., Boistel, R. & Vanhooydonck, B. 2013. Slow but tenacious: an analysis of running and gripping performance in chameleons. Journal of Experimental Biology 216, 1025-1030. pdf

Boehm, M., Collen, B., Baillie, J.E.M., Bowles, P., Chanson, J., Cox, N., Hammerson, G., Hoffmann, M., Livingstone, S.R., Ram, M., Rhodin, A.G.J., Stuart, S.N., van Dijk, P.P., Young, B.E., Afuang, L.E., Aghasyan, A., Garcia, A., Aguilar, C., Ajtic, R., Akarsu, F., Alencar, L.R.V., Allison, A., Ananjeva, N., Anderson, S., Andren, C., Ariano-Sanchez, D., Arredondo, J.C., Auliya, M., Austin, C.C., Avci, A., Baker, P.J., Barreto-Lima, A.F., Barrio-Amoros, C.L., Basu, D., Bates, M.F., Batistella, A., Bauer, A., Bennett, D., Boehme, W., Broadley, D., Brown, R., Burgess, J., Captain, A., Carreira, S., Castaneda, M.d.R., Castro, F., Catenazzi, A., Cedeno-Vazquez, J.R., Chapple, D.G., Cheylan, M., Cisneros-Heredia, D.F., Cogalniceanu, D., Cogger, H., Corti, C., Costa, G.C., Couper, P.J., Courtney, T., Crnobrnja-Isailovic, J., Crochet, P.-A., Crother, B., Cruz, F., Daltry, J.C., Daniels, R.I.R., Das, I., de Silva, A., Diesmos, A.C., Dirksen, L., Doan, T.M., Dodd, C.K., Jr., Doody, J.S., Dorcas, M.E., de Barros Filho, J.D., Egan, V.T., El Mouden, E.H., Embert, D., Espinoza, R.E., Fallabrino, A., Feng, X., Feng, Z.-J., Fitzgerald, L., Flores-Villela, O., Franca, F.G.R., Frost, D., Gadsden, H., Gamble, T., Ganesh, S.R., Garcia, M.A., Garcia-Perez, J.E., Gatus, J., Gaulke, M., Geniez, P., Georges, A., Gerlach, J., Goldberg, S., Gonzalez, J.-C.T., Gower, D.J., Grant, T., Greenbaum, E., Grieco, C., Guo, P., Hamilton, A.M., Hare, K., Hedges, S.B., Heideman, N., Hilton-Taylor, C., Hitchmough, R., Hollingsworth, B., Hutchinson, M., Ineich, I., Iverson, J., Jaksic, F.M., Jenkins, R., Joger, U., Jose, R., Kaska, Y., Kaya, U., Keogh, J.S., Koehler, G., Kuchling, G., Kumlutas, Y., Kwet, A., La Marca, E., Lamar, W., Lane, A., Lardner, B., Latta, C., Latta, G., Lau, M., Lavin, P., Lawson, D., LeBreton, M., Lehr, E., Limpus, D., Lipczynski, N., Lobo, A.S., Lopez-Luna, M.A., Luiselli, L., Lukoschek, V., Lundberg, M., Lymberakis, P., Macey, R., Magnusson, W.E., Mahler, D.L., Malhotra, A., Mariaux, J., Maritz, B., Marques, O.A.V., Marquez, R., Martins, M., Masterson, G., Mateo, J.A., Mathew, R., Mathews, N., Mayer, G., McCranie, J.R., Measey, G.J., Mendoza-Quijano, F., Menegon, M., Metrailler, S., Milton, D.A., Montgomery, C., Morato, S.A.A., Mott, T., Munoz-Alonso, A., Murphy, J., Nguyen, T.Q., Nilson, G., Nogueira, C., Nunez, H., Orlov, N., Ota, H., Ottenwalder, J., Papenfuss, T., Pasachnik, S., Passos, P., Pauwels, O.S.G., Perez-Buitrago, N., Perez-Mellado, V., Pianka, E.R., Pleguezuelos, J., Pollock, C., Ponce-Campos, P., Powell, R., Pupin, F., Quintero Diaz, G.E., Radder, R., Ramer, J., Rasmussen, A.R., Raxworthy, C., Reynolds, R., Richman, N., Rico, E.L., Riservato, E., Rivas, G., da Rocha, P.L.B., Roedel, M.-O., Rodriguez Schettino, L., Roosenburg, W.M., Ross, J.P., Sadek, R., Sanders, K., Santos-Barrera, G., Schleich, H.H., Schmidt, B.R., Schmitz, A., Sharifi, M., Shea, G., Shi, H.-T., Shine, R., Sindaco, R., Slimani, T., Somaweera, R., Spawls, S., Stafford, P., Stuebing, R., Sweet, S., Sy, E., Temple, H.J., Tognelli, M.F., Tolley, K., Tolson, P.J., Tuniyev, B., Tuniyev, S., Uzum, N., van Buurt, G., Van Sluys, M., Velasco, A., Vences, M., Vesely, M., Vinke, S., Vinke, T., Vogel, G., Vogrin, M., Vogt, R.C., Wearn, O.R., Werner, Y.L., Whiting, M.J., Wiewandt, T., Wilkinson, J., Wilson, B., Wren, S., Zamin, T., Zhou, K. & Zug, G. (2013) The conservation status of the world's reptiles. Biological Conservation, 157, 372-385. pdf

Schreiner, C., Rödder, D. & Measey, G.J. 2013. Using models to test Poynton's predictions. African Journal of Herpetology 62, 49-62. pdf

Edwards, S., Branch, W.R., Vanhooydonck, B., Herrel, A., Measey, G.J. & Tolley, K.A. 2013. Taxonomic adjustments in the systematics of southern African lacertid lizards (Sauria: Lacertidae). Zootaxa 3669, 101-114. pdf

Serclerat, H., Measey, J., Exbrayat, J.-M., & Chevalier, C. 2013. Detection histochimique des glucides dans l’ovaire d’un amphibien gymnophione: Boulengerula boulengeri pendant la periode de reproduction. Revue Française d'Histotechnologie 26 (1), 89-98. pdf

Raquet, M., Measey, J. & Exbrayat, J.-M. 2013. Mise en évidence des récepteurs des œstrogènes α et β dans les ovaires de Boulengerula taitanus amphibien gymnophione. Revue Française d'Histotechnologie 26 (1), 99-105. pdf

Measey, G.J. & Tolley, K.A. 2013. A molecular phylogeny for sub-Saharan amphisbaenians. African Journal of Herpetology 62, 100-109. pdf

Carne, L. & Measey, G.J. 2013. Chameleons on the cruise: seasonal differences in prey choice of two dwarf chameleons. Herpetological Journal 23: 221-227. pdf

Fogell, D.J., Tolley, K.A. & Measey, G.J. 2013. Mind the gaps: Investigating the cause of the current range disjunction in the Cape Platanna, Xenopus gilli (Anura: Pipidae). Peer J 1, e166.pdf  


Mokhatla, M.M., Measey, G.J., Chimimba, C.T., & van Rensburg, B.J. 2012. A biogeographical assessment of anthropogenic threats to areas where different frog breeding groups occur in South Africa: implications for anuran conservation. Diversity and Distributions 18, 470-480. pdf

Measey, G.J., Rödder, D., Green, S.L., Kobayashi, R., Lillo, F., Lobos, G., Rebelo, R. & Thirion, J.M. 2012. Ongoing invasions of the African clawed frog, Xenopus laevis: a global review. Biological Invasions 14, 2255-2270. pdf

Measey, G.J., Muchai, V., & Spawls, S. 2012. Rediscovery of Boulengerula denhardti Nieden 1912 (Amphibia:Gymnophiona: Caeciliidae) in Meru County, Kenya. African Zoology 47, 187-191. pdf

Herrel, A., Measey, G.J., Vanhooydonck, B., & Tolley, K.A. 2012. Got It Clipped? The Effect of Tail Clipping on Tail Gripping Performance in Chameleons. Journal of Herpetology 46, 91-93. pdf

Herrel, A., & Measey, G.J. 2012. Feeding Underground: Kinematics of Feeding in Caecilians. Journal of Experimental Zoology Part a-Ecological Genetics and Physiology 317A, 533-539. pdf

Edwards, S., Vanhooydonck, B., Herrel, A., Measey, G.J., & Tolley, K.A. 2012. Convergent evolution associated with habitat decouples phenotype from phylogeny in a clade of lizards. PloSone 7, e51636. pdf

Conradie, W., Branch, W.R., Measey, G.J., & Tolley, K.A. 2012. A new species of Hyperolius Rapp, 1842 (Anura: Hyperoliidae) from the Serra da Chela mountains, south-western Angola. Zootaxa, 1-17. pdf

Conradie, W., Measey, G.J., Branch, W.R., & Tolley, K.A. 2012. Revised phylogeny of African sand lizards (Pedioplanis), with the description of two new species from south-western Angola. African Journal of Herpetology 61, 91-112. pdf


Tolley, K.A., Tilbury, C.R., Measey, G.J., Menegon, M., Branch, W.R. & Matthee, C.A. 2011. Ancient forest fragmentation or recent radiation? Testing refugial speciation models in chameleons within an African biodiversity hotspot. Journal of Biogeography 38, 1748-1760. pdf

Measey, G.J. 2011. The past, present and future of African herpetology. African Journal of Herpetology 60. 89-100. pdf

Measey, G.J., & Tolley, K.A. 2011. Sequential fragmentation of Pleistocene forests in an East Africa biodiversity hotspot: chameleons as a model to track forest history. PloSone 6, e26606. pdf

Measey, G.J., & Tolley, K.A. 2011. Investigating the cause of the disjunct distribution of Amietophrynus pantherinus, the Endangered South African western leopard toad. Conservation Genetics 12, 61-70. pdf

Measey, G.J., Rebelo, A.D., Herrel, A., Vanhooydonck, B., & Tolley, K.A. 2011. Diet, morphology and performance in two chameleon morphs: do harder bites equate with harder prey? Journal of Zoology 285, 247-255. pdf

Measey, G.J., Hinkel, H.H., Dumbo, B., & Fischer, E. 2011. Rediscovery of Boulengerula fischeri, with notes on its morphology and habitat. African Journal of Herpetology 60, 47-59. pdf

Raquet, M., J. Measey & J.M. Exbrayat. 2011. Premières observations histologiques de l’oviducte de Boulengerula taitana, Loveridge, 1935, Amphibien Gymnophione. Rev Fr Histotechnol. 24: 29-38. pdf

Serclerat, H., Measey J., Exbrayat, J.-M., Chevalier, C. 2011. Etude histologique des ovaires d’un amphibian Gymnophione Boulengerula boulengeri pendant la periode de reproduction Rev. Fr. Histotechnol. 24: 39-45. pdf

Loader, S.P., Wilkinson, M., Cotton, J., Measey, G.J., Menegon, M., Howell, K.M., Muller, H. & Gower, D.J. 2011. Molecular phylogenetics of Boulengerula (Amphibia: Gymnophiona: Caeciliidae) and implications for taxonomy, biogeography and conservation. Herpetological Journal 21, 5-16. pdf

Herrel, A., Measey, G.J., Vanhooydonck, B., & Tolley, K.A. 2011. Functional consequences of morphological differentiation between populations of the Cape Dwarf Chameleon (Bradypodion pumilum). Biological Journal of the Linnean Society 104, 692-700. pdf


Bates, M.F., Pietersen, D.W., & Measey, G.J. 2010. New amphisbaenian records for the Northern Cape, South Africa. Navorsingevan die Nasionale Museum Bloemfontein 26, 61-72. pdf

Tolley, K.A., Raw, R.N.V., Altwegg, R., & Measey, G.J. 2010. Chameleons on the move: survival and movement of the Cape dwarf chameleon, Bradypodion pumilum, within a fragmented urban habitat. African Zoology  45, 99-106. pdf

Tolley, K.A., De Villiers, A.L., Cherry, M.I., & Measey, G.J. 2010. Isolation and high genetic diversity in dwarf mountain toads (Capensibufo) from South Africa. Biological Journal of the Linnean Society 100, 822-834.pdf

Malonza, P.K., Lotters, S. & Measey, G.K. 2010. The montane forest associated amphibian species of Taita Hills, Kenya. Journal of East African Natural History 99. 47-63. pdf

Herrel, A., & Measey, G.J. 2010. The kinematics of locomotion in caecilians: effects of substrate and body shape. Journal of Experimental Zoology Part a-Ecological Genetics and Physiology 313A, 301-309. pdf

Bwong, B.A., & Measey, G.J. 2010. Diet composition of Xenopus borealis in Taita Hills: effects of habitat and predator size. African Journal of Ecology 48, 299-303. pdf


Wollenberg, K.C., & Measey, G.J. 2009. Why colour in subterranean vertebrates? Exploring the evolution of colour patterns in caecilian amphibians. Journal of Evolutionary Biology 22, 1046-1056. pdf

Ngwava, J.M., Malonza, P.K., & Measey, G.J. 2009. Observations on the breeding behaviour of the Taita dwarf toad Mertensophryne taitana on Mt. Mbololo, Taita Hills, Kenya. African Journal of Herpetology 58, 44-49. pdf

Measey, G.J., Hopkins, K., & Tolley, K.A. 2009. Morphology, ornaments and performance in two chameleon ecomorphs: is the casque bigger than the bite? Zoology 112, 217-226. pdf

Measey, G.J., Armstrong, A.J., & Hanekom, C. 2009. Subterranean herpetofauna show a decline after 34 years inNdumu Game Reserve, South Africa. Oryx 43, 284-287. pdf

Loader, S.P., Measey, G.J., De Sa, R.O., & Malonza, P.K. 2009. A new Brevicipitid species (Brevicipitidae: Callulina) from the fragmented forests of the Taita Hills, Kenya. Zootaxa 2123, 55-68. pdf

Blackburn, D.C., & Measey, G.J. 2009. Dispersal to or from an African biodiversity hotspot? Molecular Ecology  18, 1904-1915. pdf


Measey, G.J., Smita, M., Beyo, R.S., & Oommen, O.V. 2008. Year-round spermatogenic activity in an oviparous subterranean caecilian, Boulengerula taitanus Loveridge 1935 (Amphibia Gymnophiona Caeciliidae). Tropical Zoology 21, 109-122. pdf


Tolley, K.A., & Measey, G.J. 2007. Chameleons and vineyards in the Western Cape of South Africa: Is automated grape harvesting a threat to the Cape Dwarf Chameleon (Bradypodion pumilum)? African Journal of Herpetology 56, 85-89. pdf

Measey, G.J., Vences, M., Drewes, R.C., Chiari, Y., Martin, M. & Bourles, B. 2007. Freshwater paths across the ocean: molecular phylogeny of the frog Ptychadena newtoni gives insights into amphibian colonization of oceanic islands. Journal of Biogeography 34, 7-20. pdf

Measey, G.J., Galbusera, P., Breyne, P., & Matthysen, E. 2007. Gene flow in a direct-developing, leaf litter frog between isolated mountains in the Taita Hills, Kenya. Conservation Genetics 8, 1177-1188. pdf



Raquet, M., Measey, G.J., & Exbrayat, J.M. 2006. Premières observations histologiques de l'ovaire deBoulangerulataitanus Loveridge, 1935, amphibien gymnophione. Revue Française d'Histotechnologie 19, 9-15.

Measey, G.J., & Van Dongen, S. 2006. Bergmann's rule and the terrestrial caecilian Schistometopum thomense  (Amphibia : Gymnophiona: Caeciiiidae). Evolutionary Ecology Research 8, 1049-1059.

Measey, G.J., Mejissa, J., & Muller, H. 2006. Notes on the distribution and abundance of the caecilian Boulengerula uluguruensis (Amphibia: Gymnophiona: Caeciliidae) in the Uluguru Mountains, Tanzania. African Journal of Ecology 44, 6-13. pdf

Measey, G.J., & Herrel, A. 2006. Rotational feeding in caecilians: putting a spin on the evolution of cranial design. Biology Letters 2, 485-487. pdf

Measey, G.J., & Barot, S. 2006. Evidence of seasonal migration in a tropical subterranean vertebrate. Journal of Zoology 269, 29-37. pdf

Measey, G.J. 2006. Surveying biodiversity of soil herpetofauna: towards a standard quantitative methodology.European Journal of Soil Biology 42, S103-S110. pdf

Fouquet, A., & Measey, G.J. 2006. Plotting the course of an African clawed frog invasion in Western France. Animal Biology 56, 95-102. pdf

Decaens, T., Jimenez, J.J., Gioia, C., Measey, G.J., & Lavelle, P. 2006. The values of soil animals for conservation biology. European Journal of Soil Biology 42, S23-S38. pdf


Muller, H., Measey, G.J., Loader, S.P., & Malonza, P.K. 2005. A new species of Boulengerula Tornier (Amphibia: Gymnophiona: Caeciliidae) from an isolated mountain block of the Taita Hills, Kenya. Zootaxa 1004, 37-50. pdf

Muller, H., Measey, G.J., & Malonza, P.K. 2005. Tadpole of Bufo taitanus (Anura: Bufonidae) with notes on its systematic significance and life history. Journal of Herpetology 39, 138-141. pdf

Measey, G.J., & Royero, R. 2005. An examination of Pipa parva (Anura: Pipidae) from native and invasive populations in Venezuela. Herpetological Journal 15, 291-294.

Measey, G.J., & Gower, D.J. 2005. Externally measured condition versus internal organ mass in the caecilian Gegeneophis ramaswamii (Amphibia : Gymnophiona : Caeciliidae). Zoological Science 22, 445-452. pdf

Malonza, P.K., & Measey, G.J. 2005. Life history of an African caecilian: Boulengerula taitanus Loveridge 1935 (Amphibia Gymnophiona Caeciilidae). Tropical Zoology 18, 49-66. pdf


Measey, G.J., Gower, D.J., Oommen, O.V., & Wilkinson, M. 2004. A subterranean generalist predator: diet of the soil-dwelling caecilian Gegeneophisramaswamii (Amphibia; Gymnophiona; Caeciliidae) in southern India. Comptes Rendus Biologies 327, 65-76. pdf

Measey, G.J. 2004. Are caecilians rare? An East African perspective. Journal of East African Natural History 93, 1-21. pdf

Gaborieau, O., & Measey, G.J. 2004. Termitivore or detritivore? A quantitative investigation into the diet of the East African caecilian Boulengerula taitanus (Amphibia: Gymnophiona: Caeciliide). Animal Biology 54, 45-56. pdf

Deletre, M., & Measey, G.J. 2004. Sexual selection vs ecological causation in a sexually dimorphic caecilian, Schistometopum thomense (Amphibia Gymnophiona Caeciliidae). Ethology Ecology & Evolution 16, 243-253. pdf


Measey, G.J., Silva, J., & Di-Bernardo, 2003. M. Testing for repeatability in measurements of length and mass in Chthonerpeton indistinctum (Amphibia: Gymnophiona), including a novel method of calculating total length of live caecilians. Herpetological Review 34, 35-39. pdf

Measey, G.J., Gower, D.J., Oommen, O.V., & Wilkinson, M. 2003. Quantitative surveying of endogeic limbless vertebrates - a case study of Gegeneophis ramaswamii (Amphibia : Gymnophiona : Caeciliidae) in southern India. Applied Soil Ecology 23, 43-53. pdf

Measey, G.J., Gower, D.J., Oommen, O.V., & Wilkinson, M. 2004. A mark-recapture study of the caecilian amphibian Gegeneophis ramaswamii (Amphibia : Gymnophiona : Caecillidae) in southern India. Journal of Zoology 261, 129-133. pdf

Measey, G.J., & Di-Bernardo, M. 2003. Estimating juvenile abundance in a population of the semiaquatic caecilian, Chthonerpeton indistinctum (Amphibia : Gymnophiona : Typhlonectidae), in southern Brazil.Journal of Herpetology 37, 371-373. pdf

Measey, G.J., & Channing, A. 2003. Phylogeography of the genus Xenopus in southern Africa. Amphibia-Reptilia  24, 321-330. pdf


Presswell, B., Gower, D.J., Oomen, O.V., Measey, G.J., & Wilkinson, M. 2002. Scolecophidian snakes in the diets of south Asian caecilian amphibians. Herpetological Journal 12, 123-126. pdf

Lobos, G., & Measey, G.J. 2002. Invasive populations of Xenopus laevis (Daudin) in Chile. Herpetological Journal12, 163-168. pdf


Measey, G.J., Gower, D.J., Oommen, O.V., & Wilkinson, M. 2001. Permanent marking of a fossorial caecilian, Gegeneophis ramaswamii (Amphibia: Gymnophiona: Caeciliidae). Journal of South Asian Natural History  5, 109-115. pdf

Measey, G.J. 2001. Growth and ageing of feral Xenopus laevis (Daudin) in South Wales, UK. Journal of Zoology  254, 547-555. pdf


Oommen, O.V., Measey, G.J., Gower, D.J., & Wilkinson, M. 2000. Distribution and abundance of the caecilian Gegeneophis ramaswamii (Amphibia : Gymnophiona) in southern Kerala. Current Science 79, 1386-1389. pdf


Moss, B., Kornijow, R., & Measey, G.J. 1998. The effects of nymphaeid (Nuphar lutea) density and predation by perch (Perca fluviatilis) on the zooplankton communities in a shallow lake. Freshwater Biology 39, 689-697. pdf

Measey, G.J., & Wilkinson, M. 1998. Lines of arrested growth in the caecilian, Typhlonectes natans (Amphibia: Gymnophiona). Amphibia-Reptilia 19, 91-95. pdf

Measey, G.J., & Tinsley, R.C. 1998. Feral Xenopus laevis in South Wales. Herpetological Journal 8, 23-27. pdf

Measey, G.J. 1998. Terrestrial prey capture in Xenopus laevis. Copeia, 787-791. pdf

Measey, G.J. 1998. Diet of feral Xenopus laevis (Daudin) in South Wales, UK. Journal of Zoology 246, 287-298. pdf


Measey, G.J., & Tinsley, R.C. 1997. Mating behavior of Xenopus wittei (Anura: Pipidae). Copeia  1997, 601-609. pdf